Paul N. Pearson (a), Eleanor H. John (a), Jamie D. Wilson (a) & Andy Ridgwell (c)
(a) School of Earth and Ocean Sciences, Cardiff University, Park Place, Cardiff CF10 3AT, UK. E-mail: pearsonp@cardiff.ac.uk
(b) School of Geographical Sciences, University of Bristol, University Road, Clifton, Bristol BS8 1SS, UK.